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To our knowledge, only few studies have focused on the crack surface itself in fossil bone54,55,56. These analyses have been limited to a superficial characterisation of the crack surfaces; they have not, however, quantified the relative thickness of the crack surfaces. Furthermore, no studies have been published about the mineral composition of the crack surfaces. Therefore, we decided to focus on the crack surfaces in the hyoid arch of the juvenile indeterminate hadrosaur Tenontosaurus dossi from the Morrison Formation (USA) (ca. 15 years; see figure 31.5 for a schematic reconstruction). The specimen was excavated from the collection of the Museum of Western Colorado (Casper, Wyoming, USA), curated by D.S. and L.R. There are two crack surfaces visible on the left mandible (figure 31.5A,B). To quantify the crack thickness, we used a chemical-mechanical polishing and a focused ion beam (FIB) milling technique53. The sample of the two crack surfaces was prepared for FIB milling using thin sections of 5-μm thickness.
The cracks generated by the LAGs may be induced by the resorptive conditions at the site, and in consequence, the “cracked” surfaces may contain a mixture of semi-decomposed periosteal material (matrix) as well as (local) calcified periosteal material (figure 31.4A). The calcified periosteum and the original bone surface may coexist side-by-side but always in close contact (figure 31.4B). What has been established for modern animals, the crack surfaces as well as the underlying bone tissue may have been further amplified in highly porous and poorly consolidated bone of dinosaurs. In highly porous and poorly consolidated bone (e.g., turtle leg bone), the crack surfaces may be characterized by a non-uniform composition of the material, which may contain a mixture of calcified and semi-decomposed periosteal material (figure 31.4C,D). Conversely, in bones containing an articular surface (e.g., endochondral bone of the hyoid arch, periosteal bone of ulna and radius of pterosaurs), the crack surfaces may be characterized by an organized matrix/calcified matrix interface (figure 31.4E,F).
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